Michael D. Jemings
U.S. Fish and Wildlife Service
Idaho Cooperative Fish and Wildlife Research Unit
University of Idaho
Moscow, Idaho 83843 USA
Table 1. A land and water classification system for Gap Analysis.
LEVEL 1 LEVEL 2 LEVEL 3
--Forest--Mainly Evergreen -Mainly Deciduous
Mainly Evergreen -Woodlands ~ainl~
Deciduous L~xtremel~
Xeromomphic
Mainly Evergreen -Shrublands Mainly Deciduous Extremely Xeromorophic Natural Terrestrial Cover Mainly Evergreen warf-Shrublands Mainly Deciduous -Extremely Xeromorophic Tundra
Grasslands Short
ry Salt Flats Beaches andy Areas
erennial Snowfields
,--fiverim drower
Upper
ntermittent
Natural Aquatic Cover Palustrine None
Estuarine p u b t i d a l
ntertidal
Row Crop, Pasture rchard, Horticulture Agriculture 4Confined Livestock Feeding Other Agriculture Developed Cover Residential ommerciauIndustrial
-Urban Transportation ther
Natural Temesm'al Cover Cassification: Assumptions and Definitions
Bailey is currently revising the ecoregion map of the United States, based on better information and on an improved understanding of the integrated relationships among climate, land surface, vegetation, and scale, resulting partially from work done to map ecoregions of the continents (Bailey, 1989; personal communication).
A. All references and applications are made to actual land cover rather than historic land cover or potential climax vegetation.
B. Classifications of natural systems are imprecise; there will always be zones of transition and variants, and vegetation boundaries may not be discrete.
C. The criteria used to define classification categories (e.g., tree savannah = tree canopy cover of 11-25 % over grassland) are general. They are meant as data quality objectives (DQOs, see definitions) as a means for grouping and discussing discrete cover types, from coarse categories such as grasslands to finer categories such as short-grass comniunities composed of bunch grasses.
D. In order to create useful categories across both biotic and geographic scales, it is necessary to use both subdivisions and aggregations. Subdivision begins with a whole and divides it into smaller units according to similarities. Aggregation begins with a set of individual objects and groups them according to similarities. For example, classifications at the regional or landscape level (e.g., evergreen coniferous woodlands with rounded crowns) are based on the physioenomic subdivisions of class, subclass, group, and formation. Classifications at the minimum mapping unit level (e.g., Juniperus occidentalislArtemisia tridentata) are based on a floristic aggregation of associated plant species up to the community type level (Table 2; see definitions).
E. The classification system should fit within and among existing accepted national and international land cover classification systems (e.g., UNESCO and The Nature Conservancy for mapping vegetation, U.S. Fish and Wildlife Service classification of wetlands and deepwater habitats, U.S. Geological Survey classification of land use and land cover).
F. The classification scheme is open-ended. Categories may be added to any of the hierarchical levels (e. g., class, subclass, group, formation) as long as the additions are truly an equivalent category within the given classification level.
G. The classification scheme is intended to be the super-structure upon which information can continuously be placed for meaningful analysis and interpretation. It is assumed that:
B. associated forms of vegetation, e.g.,
mixed with herbaceous plants;
C. climate, e.g., cold, drought;
D. morphology, e.g., evergreen,
deciduous.
111. for grasslands:
A. tree or shrub canopy less than 10 % for areas dominated by graminoids;
B. tree canopy or shrub canopy 11 -25 %
for areas dominated by graminoids,
e.g., tree or shrub savannah;
C. alpine and subalpine occurrence;
D. areas dominated by forbs more than 1
meter in height;
E. areas dominated by forbs less than 1
meter in height.
Formation: Categories within each group
comprised of areas in which the vegetation
similarities are based on any of the following
criteria.
I. tree size and crown shape:
A. giant forests are those more than 50
meters in height having
B. non-giant forests are those 5-50 meters in height having
11. life zone:
A. temperate lowland
B. montane
C. alpine
D. boreal
111. substrate:
A. alluvial
B. serpentine
Cover Type: A group of plant community types having the same primary dominant species and similar physiognomy; an aggregation of plant community types.
Community Type: An assemblage of plant species that interact at the same time and place, of defined species composition, and physiognomy, regardless of sera1 stage; usually named by combining the name of the species that dominates the canopy layer with the name of the species that dominates lower vegetation layers (e.g., Juniperus occidentalis /Arternisia tridentata) .
Other terms used in the classification of terrestrial vegetation for Gap Analysis and their definitions follow.
Data Quality Objectives: The accuracy and precision objectives for data representation of location and object attributes.
Dominant or Codominant: Pertaining to the plant species having the greatest relative canopy cover.
Forb: A non-graminoid herb.
Natural Community: A discrete group of plant and animal species that reoccur across a landscape.
Natural Vegetation: Vegetation that is regenerating without cultivation regardless of previous disturbance; not restricted to undisturbed climax vegetation.
Shrubs: Woody perennial plants, usually with
APPENDIX A
A KEY TO NATURAL TERRESTRIAL VEGETATION FORMATIONS'
'~rom Driscoll, R.S., D.L. Merkel, D.L. Radloff, D.E. Snyder, and J. S. Hagihara. 1984. An ecological land classification framework for the United States. U.S.D.A. Forest Service, Miscellaneous Publication 1439, Washington, D.C. p. 21.
Class: ~orestl
Subclass Group Formation
| Trees over 5 m tall forming 61 to 100 | A. | Mainly evergreen forest (canopy | 1. | Tropical rain forest (mainly broad- | a. | Tropical lowland rain forest |
| percent canopy cover | never without green foliage although | leaved evergreen trees, neither cold- | b. | Tropical submontane rain forest | ||
| some trees may shed their leaves) | nor drought--resistant) | c. | Tropical montane rain forest | |||
| d. | Tropical "subalpine" rain forest | |||||
| e. | Tropical (rain) cloud forest | |||||
| f. | Tropical alluvial rain forest | |||||
| g. h. | Tropical swamp rain forest Tropical evergreen bog forest | |||||
| (classical tropical rain forests) | ||||||
| 2. | Tropical and subtropical evergreen | a. | Tropical or subtropical evergreen | |||
| seasonal forest (mainly broad-leaved | seasonal lowland forest | |||||
| evergreen trees with some foliage | b. | Tropical or subtropical evergreen | ||||
| reduction in dry season) | seasonal submontane forest | |||||
| C. | Tropical or subtropical evergreen | |||||
| seasonal montane forest | ||||||
| Tropical or subtropical evergreen dry | ||||||
| "subalpine" forest | ||||||
| 3. | Tropical and subtropical | Tropical or subtropical | ||||
| semideciduous forest (upper canopy | semideciduous lowland forest | |||||
| trees drought deciduous) | Tropical or subtropical | |||||
| semideciduous montane or cloud | ||||||
| forest | ||||||
| 4. | Subtropical rain forest (not | Subtropical lowland rain forest | ||||
| represented in U .S.) | Subtropical submontane rain forest | |||||
| Subtropical montane rainforest | ||||||
| Subtropical "subalpine" rain forest | ||||||
| Subtropical (rain) cloud forest | ||||||
| Subtropical alluvial rain forest | ||||||
| Subtropical swamp rain forest | ||||||
| Subtropical evergreen bog forest | ||||||
| (with organic surface deposits) |
| Class: woodland1 | ||||||
|---|---|---|---|---|---|---|
| Class:Woodland | Subclass | Group | Formation | |||
| Open to 60 forest (trees over 5 m tall forming 26 percent canopy cover) | A. | Mainly evergreen woodland (canopy never without green foliage although some trees may shed their leaves) | 1 . 2. | Evergreen broad-leaved woodland Evergreen needle-leaved woodland | (No formations defined; includes evergreen oak woodlands) a. Evergreen needle-leaved woodland with rounded crowns (e.g., pine and juniper) b . Evergreen needle-leaved woodland with conical crowns (e.g., spruce in the west) c, Evergreen needle-leaved woodland with very narrow cylindro-conical crowns (e.g., some spruce in Alaska) | |
| B . | Mainly deciduous woodland (majority of trees shed foliage as result of drought or cold) | 1. | Tropical and subtropical drought- deciduous woodland (frost and freeze generally absent or moderate) | a. b . | Drought-deciduous broad-leaved lowland and submontane woodland Drought-deciduous montane (and cloud) woodland | |
| 2. | Cold-deciduous woodland, with evergreen trees (winter frost and freeze) | a. b . | Evergreen broad-leaved woodland Evergreen needle-leaved woodland | |||
| 3. | Cold-deciduous woodland, without evergreen trees (winter frost and freeze) | a. b . c. | Broad-leaved deciduous woodland. Needle-leaved deciduous woodland Mixed deciduous woodland (broad- leaved and needle-leaved) | |||
| C . | Extremely xeromorphic woodland (dry woodlands) | 1. | Scierophyllous woodland | a. | Evergreen scierophyllous-dominated xeromorphic woodland | |
| 2. | Thorn-woodland (may not be represented in U.S.) | a. b . | Mixed deciduous-evergreen thom woodland Purely deciduous thom woodland | |||
| 3. | Mainly succulent woodland | |||||
l~hekey to the vegetation element is patterned after UNESCO 1973. The key contains a few exainples of known communities in the United States.
| Class: Herbaceous Vegetationl | ||||
|---|---|---|---|---|
| C1ass:fIerbaceous Vegetation | Subclass | Group | ||
| Trees or shrubs occupy 25 percent or less of the canopy cover | A. | Tall grassland (grasses or forbs over 1 m tall) | 1. | Tall grassland with a tree layer (tree canopy cover is 11 to 25 percent (e.g., the transition from the true tall grassland prairie to the forests; a tree savannah) |
| 2. | Tall grassland with a shrub layer (shrub canopy cover is 11 to 25 percent) (e. g.. the transition from the true tall grassland prairie and shrub land; a shrub savannah) | |||
| 3. | Tall grassland with open layer of tuft plants, usually palms (specific examples unknown for U.S.) | |||
| 4. | Tall grassland without a woody layer (tree or shrub canopy cover is 0 to 10 percent) (e.g., tall grass prairie of eastern Kansas) | |||
| B. | Medium tall grassland (grasses and forbs over 0.5 to 1 m tall) | 1. | Medium tall grassland with a tree layer (tree canopy cover is 11 to 25 percent) (as in subclass tall grassland -the tree savannah) | |
| 2. | Medium tall grassland with a shrub layer (shrub canopy cover is 11 to 25 percent) (as in subclass medium tall grassland -the shrub savannah) | |||
Formation
Woody layer broad-leaved evergreen Woody layer broad-leaved semi- evergreen Woody layer broad-leaved deciduous Woody layer needle-leaved evergreen and broad-leaved deciduous Woody layer needle-leaved evergreen
Shrub layer broad-leaved evergreen Shrub layer broad-leaved semi- evergreen Shrub layer broad-leaved deciduous
Tall subtropical grassland with open groves of palms
Tall grassland consisting mainly of sod grasses Tall grassland consisting mainly of bunch grasses
Woody layer broad-leaved evergreen Woody layer broad-leaved semi- evergreen Woody layer broad-leaved deciduous Woody layer needle-leaved evergreen and broad-leaved deciduous Woody layer needle-leaved evergreen Woody layer needle-leaved evergreen and broad-leaved evergreen
Shrub layer broad-leaved evergreen Shrub layer broad-leaved semi- evergreen Shrub layer broad-leaved deciduous Shrub layer of mainly deciduous thorny shrubs Shrub layer of mainly needle-leaved evergreen
| s: Herbaceous Vegetation -Continued | ||||||
|---|---|---|---|---|---|---|
| -C1ass:Herbaceous Vegetation | Subclass | Group | Formation | |||
| 6. | Mesophytic grassland(a1pine and subalpine meadows)(e.g., bluegrass and hairgrass communities) | a. b. | Sod grass communities usually dominated by hemicrtophytes Alpine and subalpine meadows of the higher latitudes | |||
| 7. | Short grassland tundra (grass tundra of Alaska) | a. b. | Bunch forming short grassland tundra Sod forming short grassland tundra | |||
| D. | Forb-dominated vegetation (herbaceous communities with forb cover exceeding 50 percent) | 1. | Tall forbs (forbs over 1 m tall) (tall forb meadows, Utah mountains) | a. b. c. | Mainly perennial flowering forbs, and ferns Fern communities especially in humid climates Mainly annual forbs | |
| 2. | Low forbs (forbs 1 m or less tall) (Aleutian forb meadows, Alaska) | a. b. | Mainly perennial flowering forhs, and ferns Mainly annual forbs | |||
he key to the vegetation element is patterned after UNESCO 1973. The key contains a few examples of known communities in the United States.
BAILEY'S SECI~O?-
Rcparcd from Ecorcgiona of the United Srrtcr, Robcr( G. Bailey, 1976, by Michael lenninga and Joel Hall, 1993,
U.S. Fiah and Wildlife Service Gap Anrlyaia Rognm, Idaho Coopenlive Fil and Wildlife Research Unit, Univenity of Idaho, Moocow, 83843.
ILEVEL ECOREGIONS
APPENDIX C
A LIST OF STATES BY EACH OF BAILEY'S SECTION-LEVEL ECOREGIONS
| Spruce-Fir Forest (2111) | MN, WI. |
| Northern Hardwoods-Fir Forest (21 12) | MI, WI. |
| Northern Hardwoods Forest (21 13) | MI, NJ, NY, OH, PA, WI. |
| Northern Hardwoods-Spruce Forest (21 14) | CT, MA, ME, NH, NY, VT. |
| Mixed Mesophytic Forest (221 1) | AL, GA, KY, OH, PA, TN, VA, WV. |
| Beech-Maple Forest (22 12) | IN, MI, NY, OH, PA. |
| Maple-Basswood Forest + Oak Savanna (22 13) | IA, IL, MN, WI. |
| Appalachian Oak Forest (22 14) | CT, DE, GA, MA, MD, ME, NC, NH, |
| NJ, NY, OH, PA, RI, SC, TN, VA, | |
| wv. | |
| Oak-Hickory Forest (22 15) | AL, AR, IL, IN, KY, MO, MS, OK, |
| TN. | |
| Beech-Sweetgum-Magnolia-Pine-OakForest (2311) | AL, FL, GA, LA, MS, SC TX. |
| Southern Floodplain Forest (23 12) | AL, AR, LA, MO, MS, TN. |
| Southeastern Mixed Forest (2320) | AL, AR, DE, GA, LA, MD, MS, NC, |
| NJ, SC, TX,VA. | |
| Willarnette-Puget Forest (2410) | OR, WA. |
| Oak-Hickory-Bluestem Parkland (25 11) | IL, IA, IN, KS, MO, OK. |
| Oak + Bluestem Parkland (25 12) | KS, OK, TX |
| Mesquite-Buffalo Grass (252 1) | TX,OK. |
| Juniper-Oak-Mesquite (2522) | TX. |
| Mesquite-Acacia (2523) | TX. |
| Bluestem Prairie (2531) | IA, KS, MN, MO, ND, NE, OK, SD. |
| ND, NE, SD. |
California Mixed Evergreen Forest (M2414)
Silver Fir-Douglas Fir Forest (M2415)
Sierran Forest (M2610)
California Chaparral (M2620)
Grand Fir-Douglas Fir Forest (M3111)
Douglas Fir Forest (M3112)
Ponderosa Pine-Douglas Fir Forest (M3113)
Juniper-Pinyon Woodland +
Sagebrush-Saltbush Mosaic (P3 13 1)
Grarna-Galleta Steppe + Juniper-Pinyon Woodland Moasic (P3 132) Wheatgrass-Needlegrass-Sagebrush (A3 141) Sagebrush-Wheatgrass (A3 142) CA.
OR, WA.
CA, OR.
CA.
ID, OR, WA.
CO, ID, MT, UT, WY.
CO, NM, WY.
AZ, CO, UT.
AZ, CO, NM.
MT, WY.
ID, CO, UT, WY.
Terrestrial Vegetation Clmsificarion: Assumptions and Definitions
KY Oak-Hickory Forest (2215) Mixed Mesophytic Forest (221 1)
LA Southern Floodplain Forest (23 12) Southeastern Mixed Forest (2320) Beech-Sweetgum-Magnolia-Pine-Oak Forest (23 1 1)
MA Northern Harwoods-Spruce Forest (21 14) Appalachian Oak Forest (2214)
MD Southeastern Mixed Forest (2320) Appalachian Oak Forest (2214)
ME Northern Harwoods-Spruce Forest (21 14) Appalachian Oak Forest (2214)
MI Northern Hardwoods Forest (21 13) Northern Hardwoods-Fir Forest (21 12) Beech-Maple Forest (2212)
MO Oak-Hickory-Bluestem Parkland (251 1) Oak-Hickory Forest (22 15) Southern Floodplain Forest (23 12)
MN Spruce-Fir Forest (21 1 1) Maple-Basswood Forest + Oak Savanna (22 1 3) Bluestem Prairie (253 1)
MS Southern Floodplain Forest (23 12) Southeastern Mixed Forest (2320) Oak-Hickory Forest (22 15) Beech-Sweetgum-Magnolia-Pine-Oak Forest (23 1 1)
MT Grama-Needlegrass-Wheatgrass Prairie (31 11) Wheatgrass-Needlegrass Prairie (31 12) Wheatgrass-Needlegrass Sagebrush Steppe (A3141) Douglas Fir Forest (M3112) Cedar-Hemlock-Douglas Fir Forest (M2112)
NC Appalachian Oak Forest (21 14)
Southeastern Mixed Forest (2320)
ND Wheatgrass-Needlegrass Prairie (3 112) Wheatgrass-Bluestem-Needlegrass Prairie (31 12) ND Bluestem Prairie (2531)
NE Wheatgrass-Needlegrass Prairie (3 1 12) Bluestem Prairie (253 1) Wheatgrass-Bluestem-Needlegrass Prairie (2532) Bluestem-Grama Prairie (2533) Grama-Buffalo Grass Prairie (3 1 13)
NH Northern Hardwoods-Spruce Forest (21 14) Appalachian Oak Forest (2214)
NJ Appalachian Oak Forest (2214) Southeastern Mixed Forest (2320) NJ Northern Hardwoods Forest (2113)
NM Grama-Galleta Steppe + Juniper-Pinyon Woodland Moasic (P3 132) Upper Gila Mountains Forest (M3 120) Juniper-Pinyon Woodland + Sagebrush-Saltbush Mosaic (P3 131) Mexican Highlands Shrub Steppe (3140) Grama-Tobosa Desert (321 1) Tarbrush-Creosote Bush Desert (3212) Grama-Buffalo Grass Prairie (31 13)
NV Sierran Forest (M2610) Sagebrush-Wheatgrass Steppe (3 13 1) Lahontan Saltbush-Greaswood Steppe (3132) Great Basin Sagebrush Steppe (3133) Bonneville Saltbush-Greaswood Steppe (3 134) Creosote Bush Desert (3221)
NY Northern Hardwood Forest (21 13)
Appalachian Oak Forest (2214)
Beech-Maple Forest (2212)
OH Beech-Maple Forest (22 12)
Appalachian Oak Forest (2214)
Mixed Mesophytic Forest (2212)
Oak-Hickory Forest (2215)
Northern Hardwood Forest (21 13)
Temsniol Vegetation Chsification: Assumptions and Dejnitions
WY Sagebrush-Wheatgrass(A3 142) Wheatgrass-Needlegrass-Sagebrush (A3 141) Wheatgrass-Needlegrass Prairie (3 1 12) Grarna-Buffalo Grass Prairie (3 1 13) Ponderosa Pine-Douglas Fir Forest (M3 1 13) Douglas Fir Forest (M3 1 12)
Temsmmal Vegeration Clasajication: Assumptions and Defintions